All buttons have a transparent background

Hi, 
On the front page of my website: theluxurytraveller.com I have a "search" button and a "book online instantly" button.
The search button is set to default theme colors by the search widget and the other has been added using the button shortcode.
I defined the color blue in the shortcode but this has only been applied to the outline.

It seems that somewhere in my theme there is an override making the background colour of these buttons transparent.
Where can I alter this?

I've tried Theme options>colors>content  but the button settings here have no impact.


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